About the Authors

Joanne L. Edgar

j.edgar@bris.ac.uk

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Elizabeth S. Paul

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Lauren Harris

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Sarah Penturn

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Christine J. Nicol

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CJN ESP JLE. Performed the experiments: JLE LH SP. Analyzed the data: JLE. Contributed reagents/materials/analysis tools: JLE CJN ESP. Wrote the paper: JLE CJN ESP.