About the Authors

Raymond Noordam

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Steffy W. M. Jansen

Contributed equally to this work with: Steffy W. M. Jansen, Abimbola A. Akintola

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Abimbola A. Akintola

Contributed equally to this work with: Steffy W. M. Jansen, Abimbola A. Akintola

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Nicole Y. L. Oei

Affiliations Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands, Department of Radiology, Leiden University Medical Center, Leiden, the Netherlands, Leiden Institute for Brain and Cognition, Leiden, the Netherlands

Andrea B. Maier

Affiliations Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands, Netherlands Consortium of Healthy Ageing, the Netherlands

Hanno Pijl

Affiliation Department of Endocrinology, Leiden University Medical Center, Leiden, the Netherlands

P. Eline Slagboom

Affiliations Section of Molecular Epidemiology, department of Medical Statistics and Bioinformatics, Leiden University Medical Center, Leiden, the Netherlands, Netherlands Consortium of Healthy Ageing, the Netherlands

Rudi G. J. Westendorp

Affiliations Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands, Netherlands Consortium of Healthy Ageing, the Netherlands

Jeroen van der Grond

Affiliation Department of Radiology, Leiden University Medical Center, Leiden, the Netherlands

Anton J. M. de Craen

Affiliations Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands, Netherlands Consortium of Healthy Ageing, the Netherlands

Diana van Heemst

d.van_heemst@lumc.nl

Affiliations Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands, Netherlands Consortium of Healthy Ageing, the Netherlands

On behalf of the Leiden Longevity Study group

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RN SWMJ AAA DvH. Performed the experiments: RN. Analyzed the data: RN. Contributed reagents/materials/analysis tools: AJMdeC. Wrote the paper: RN SWMJ AAA DVH. Initial data collection and study design of the Leiden Longevity Study and its subcohorts: PES RGJW AJMdC ABM DvH. Data quality and advising the analyses: NYLO JvdG. Endocrinological advise: HP. Critically reading and commenting on the versions of the initial manuscript: DvH RN SWMJ AAA NYLO ABM HP PES RGJW JvdG AJMdC.