About the Authors

Faith Hall-Glenn

Contributed equally to this work with: Faith Hall-Glenn, R. Andrea De Young

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

R. Andrea De Young

Contributed equally to this work with: Faith Hall-Glenn, R. Andrea De Young

Current address: Amgen, Thousand Oaks, California, United States of America

Affiliation Department of Orthopaedic Surgery, University of California Los Angeles, Los Angeles, California, United States of America

Bau-Lin Huang

Current address: Center for Cancer Research, National Cancer Institute, Frederick, Maryland, United States of America

Affiliation Department of Oral Biology, University of California Los Angeles, Los Angeles, California, United States of America

Ben van Handel

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Jennifer J. Hofmann

Current address: Karolinska Institutet, Stockholm, Sweden

Affiliations Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Molecular Biology Institute, University of California Los Angeles, Los Angeles, California, United States of America

Tom T. Chen

Current address: South San Francisco, California, United States of America

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Aaron Choi

Affiliation Department of Orthopaedic Surgery, University of California Los Angeles, Los Angeles, California, United States of America

Jessica R. Ong

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Paul D. Benya

Affiliation Department of Orthopaedic Surgery, University of California Los Angeles, Los Angeles, California, United States of America

Hanna Mikkola

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

M. Luisa Iruela-Arispe

Affiliations Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Molecular Biology Institute, University of California Los Angeles, Los Angeles, California, United States of America, Jonsson Comprehensive Cancer Center, University of California Los Angeles, Los Angeles, California, United States of America

Karen M. Lyons

klyons@mednet.ucla.edu

Affiliations Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Department of Orthopaedic Surgery, University of California Los Angeles, Los Angeles, California, United States of America, Molecular Biology Institute, University of California Los Angeles, Los Angeles, California, United States of America, Jonsson Comprehensive Cancer Center, University of California Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KML MLI-A HM. Performed the experiments: FH-G RAD B-LH BvH JJH TTC JRO AC PDB. Analyzed the data: FH-G MLI-A HM PDB KML. Contributed reagents/materials/analysis tools: PDB HM MLI-A. Wrote the paper: FH-G KML.