About the Authors

David Schley

David.Schley@iah.ac.uk

Affiliation Institute for Animal Health, Pirbright, Woking, United Kingdom

Reiko J. Tanaka

Affiliation Department of Bioengineering, Imperial College London, London, United Kingdom

Kritsada Leungchavaphongse

Affiliation Department of Bioengineering, Imperial College London, London, United Kingdom

Vahid Shahrezaei

Affiliation Department of Mathematics, Imperial College London, London, United Kingdom

John Ward

Affiliation Department of Mathematical Sciences, Loughborough University, Leicestershire, United Kingdom

Clare Grant

Affiliation Institute for Animal Health, Pirbright, Woking, United Kingdom

Bryan Charleston

Affiliation Institute for Animal Health, Pirbright, Woking, United Kingdom

Christopher J. Rhodes

Affiliation Institute for Mathematical Sciences, Imperial College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DS BC CG JW. Performed the experiments: RJT VS KL CG DS. Analyzed the data: KL RJT DS CJR JW. Contributed reagents/materials/analysis tools: VS JW RJT CJR DS. Wrote the paper: DS CJR RJT JW.