About the Authors

Rosie Woodroffe

rosie.woodroffe@ioz.ac.uk

Affiliations Institute of Zoology, Regent's Park, London, United Kingdom, Wildlife Health Center, University of California Davis, Davis, California, United States of America

Katherine C. Prager

Affiliation Wildlife Health Center, University of California Davis, Davis, California, United States of America

Linda Munson

† Deceased.

Affiliations Wildlife Health Center, University of California Davis, Davis, California, United States of America, Department of Pathology, Microbiology and Immunology, University of California Davis, Davis, California, United States of America

Patricia A. Conrad

Affiliations Wildlife Health Center, University of California Davis, Davis, California, United States of America, Department of Pathology, Microbiology and Immunology, University of California Davis, Davis, California, United States of America

Edward J. Dubovi

Affiliation Animal Health Diagnostic Center, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Jonna A. K. Mazet

Affiliation Wildlife Health Center, University of California Davis, Davis, California, United States of America

Competing Interests

This work was partly funded by conservation funds associated with commercial operations (Denver Zoo, Columbus Zoo, Philadelphia Zoo). This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RW KCP LM JAKM. Performed the experiments: RW KCP PAC EJD. Analyzed the data: RW. Contributed reagents/materials/analysis tools: PAC EJD. Wrote the paper: RW KCP PAC EJD JAKM.