About the Authors

Karen A. Alexander

Karen.Alexander@sams.ac.uk

Affiliation Scottish Association for Marine Science, Scottish Marine Institute, Oban, Argyll, United Kingdom

Ron Janssen

Affiliation Institute for Environmental Studies, Vrije Universiteit, Amsterdam, The Netherlands

Gustavo Arciniegas

Affiliation Institute for Environmental Studies, Vrije Universiteit, Amsterdam, The Netherlands

Timothy G. O'Higgins

Affiliation Scottish Association for Marine Science, Scottish Marine Institute, Oban, Argyll, United Kingdom

Tessa Eikelboom

Affiliation Institute for Environmental Studies, Vrije Universiteit, Amsterdam, The Netherlands

Thomas A. Wilding

Affiliation Scottish Association for Marine Science, Scottish Marine Institute, Oban, Argyll, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KAA RJ GA TGO TAW. Performed the experiments: KAA RJ GA TGO TE. Analyzed the data: KAA. Contributed reagents/materials/analysis tools: RJ GA TE. Wrote the paper: KAA RJ GA TGO TAW.