About the Authors

Heather M. Fritz

Contributed equally to this work with: Heather M. Fritz, Paul W. Bowyer

Affiliation Department of Pathology, Microbiology and Immunology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Paul W. Bowyer

Contributed equally to this work with: Heather M. Fritz, Paul W. Bowyer

Affiliations Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America, Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Matthew Bogyo

Affiliations Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America, Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Patricia A. Conrad

paconrad@ucdavis.edu (PC); john.boothroyd@stanford.edu (JB)

Affiliation Department of Pathology, Microbiology and Immunology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

John C. Boothroyd

paconrad@ucdavis.edu (PC); john.boothroyd@stanford.edu (JB)

Affiliation Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HF PB PC JB. Performed the experiments: HF PB. Analyzed the data: HF PB. Contributed reagents/materials/analysis tools: MB JB PC. Wrote the paper: HF PB PC JB.