About the Authors

Takafumi Hasegawa

thasegawa@med.tohoku.ac.jp

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Masatoshi Konno

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Toru Baba

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Naoto Sugeno

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Akio Kikuchi

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Michiko Kobayashi

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Emiko Miura

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Nobuyuki Tanaka

Affiliation Division of Cancer Biology and Therapeutics, Miyagi Cancer Center Research Institute, Natori, Miyagi, Japan

Keiichi Tamai

Affiliation Division of Cancer Biology and Therapeutics, Miyagi Cancer Center Research Institute, Natori, Miyagi, Japan

Katsutoshi Furukawa

Affiliation Department of Geriatrics and Gerontology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Miyagi, Japan

Hiroyuki Arai

Affiliation Department of Geriatrics and Gerontology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Miyagi, Japan

Fumiaki Mori

Affiliation Department of Neuropathology, Institute of Brain Science, Hirosaki University School of Medicine, Hirosaki, Aomori, Japan

Koichi Wakabayashi

Affiliation Department of Neuropathology, Institute of Brain Science, Hirosaki University School of Medicine, Hirosaki, Aomori, Japan

Masashi Aoki

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Yasuto Itoyama

Affiliations Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan, National Center Hospital for Mental, Nervous, and Muscular Disorders, National Center of Neurology and Psychiatry, Kodaira, Tokyo, Japan

Atsushi Takeda

Affiliation Division of Neurology, Department of Neuroscience & Sensory Organs, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TH AT. Performed the experiments: TH M. Konno TB NS AK M. Kobayashi EM FM KW. Analyzed the data: TH M. Konno TB. Contributed reagents/materials/analysis tools: NT KT KF HA KW MA YI. Wrote the paper: TH.