About the Authors

Nils Schoof

Affiliations Section of Epidemiology and Biostatistics, Leeds Institute of Molecular Medicine, University of Leeds, Leeds, United Kingdom, Department of Medical Epidemiology and Biostatistics, Karolinska Institute, Stockholm, Sweden

Mark M. Iles

Affiliation Section of Epidemiology and Biostatistics, Leeds Institute of Molecular Medicine, University of Leeds, Leeds, United Kingdom

D. Timothy Bishop

Affiliation Section of Epidemiology and Biostatistics, Leeds Institute of Molecular Medicine, University of Leeds, Leeds, United Kingdom

Julia A. Newton-Bishop

Affiliation Section of Epidemiology and Biostatistics, Leeds Institute of Molecular Medicine, University of Leeds, Leeds, United Kingdom

Jennifer H. Barrett

J.H.Barrett@leeds.ac.uk

Affiliation Section of Epidemiology and Biostatistics, Leeds Institute of Molecular Medicine, University of Leeds, Leeds, United Kingdom

GenoMEL consortium

Affiliation The Melanoma Genetics Consortium (GenoMEL)

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JHB NS MMI JANB DTB. Performed the experiments: NS . Analyzed the data: NS JHB. Wrote the paper: NS JHB. Designed study and provided samples generating the original GWA data on which the analysis is based: GenoMEL consortium.