About the Authors

Sofia I. Gabriel

Affiliations Department of Biology, University of York, York, United Kingdom, CESAM–Centre for Environmental and Marine Studies, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, Lisbon, Portugal

Mark I. Stevens

Affiliation South Australian Museum, and School of Earth and Environmental Sciences, University of Adelaide, Adelaide, Australia

Maria da Luz Mathias

Affiliation CESAM–Centre for Environmental and Marine Studies, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, Lisbon, Portugal

Jeremy B. Searle

jeremy.searle@cornell.edu

Affiliations Department of Biology, University of York, York, United Kingdom, Department of Ecology and Evolutionary Biology, Cornell University, Corson Hall, Ithaca, New York, United States of America

Competing Interests

The authors have read the journal's policy and have the following conflicts: Part of this work was carried out by using the resources of the Computational Biology Service Unit from Cornell University which is partially funded by Microsoft Corporation. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: SIG MIS MLM JBS. Performed the experiments: SIG. Analyzed the data: SIG. Contributed reagents/materials/analysis tools: SIG MIS. Wrote the paper: SIG MIS MLM JBS.