About the Authors

Paola Neri

Affiliation Microbiology, Department of Biopharmaceutical Sciences, Gifu Pharmaceutical University, Gifu, Japan

Shunji Tokoro

Affiliation Microbiology, Department of Biopharmaceutical Sciences, Gifu Pharmaceutical University, Gifu, Japan

Ryo Kobayashi

Affiliation Microbiology, Department of Biopharmaceutical Sciences, Gifu Pharmaceutical University, Gifu, Japan

Tsuyoshi Sugiyama

Affiliation Microbiology, Department of Biopharmaceutical Sciences, Gifu Pharmaceutical University, Gifu, Japan

Kouji Umeda

Current address: Immunology Research Institute, EW Nutrition Japan K.K., Gifu, Japan

Affiliation Immunology Research Institute, GHEN Corporation, Gifu, Japan

Takeshi Shimizu

Affiliation Department of Molecular Infectiology, Graduate School of Medicine, Chiba University, Chiba, Japan

Takao Tsuji

Affiliation Department of Microbiology, School of Medicine, Fujita Health University, Aichi, Japan

Yoshikatsu Kodama

Current address: Immunology Research Institute, EW Nutrition Japan K.K., Gifu, Japan

Affiliation Immunology Research Institute, GHEN Corporation, Gifu, Japan

Keiji Oguma

Affiliation Department of Bacteriology, Graduate School of Medicine, Dentistry and Pharmaceutical Sciences, Okayama University, Okayama, Japan

Hiroshi Mori

morih@gifu-pu.ac.jp

Affiliation Microbiology, Department of Biopharmaceutical Sciences, Gifu Pharmaceutical University, Gifu, Japan

Competing Interests

Contributing authors K. Umeda and Y. Kodama are full-time employees of Immunology research Institute, GHEN Corporation. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors. The other authors declare that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YK T. Sugiyama KO HM. Performed the experiments: PN RK ST KU HM. Analyzed the data: PN T. Sugiyama HM. Contributed reagents/materials/analysis tools: T. Shimizu TT. Wrote the paper: PN HM.