About the Authors

Scott A. Houck

Contributed equally to this work with: Scott A. Houck, Andrew Landsbury

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington, United States of America

Andrew Landsbury

Contributed equally to this work with: Scott A. Houck, Andrew Landsbury

Affiliation School of Biological and Biomedical Sciences, University of Durham, Durham, United Kingdom

John I. Clark

r.a.quinlan@dur.ac.uk (RAQ); clarkji@u.washington.edu (JIC)

Affiliations Department of Biological Structure, University of Washington, Seattle, Washington, United States of America, Department of Ophthalmology, University of Washington, Seattle, Washington, United States of America

Roy A. Quinlan

r.a.quinlan@dur.ac.uk (RAQ); clarkji@u.washington.edu (JIC)

Affiliations School of Biological and Biomedical Sciences, University of Durham, Durham, United Kingdom, Biophysical Sciences Institute, University of Durham, Durham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SH AL JIC RAQ. Performed the experiments: SH AL. Analyzed the data: SH AL JIC RAQ. Contributed reagents/materials/analysis tools: SH AL JIC RAQ. Wrote the paper: SH AL JIC RAQ.