About the Authors

Neeltje van Doremalen

Affiliations Department of Virology, Division of Infectious Disease, St. Mary's Campus, Imperial College, London, United Kingdom, Virus Reference Department, Health Protection Agency Microbiology Services Colindale, London, United Kingdom

Holly Shelton

Affiliation Department of Virology, Division of Infectious Disease, St. Mary's Campus, Imperial College, London, United Kingdom

Kim L. Roberts

Affiliation Department of Virology, Division of Infectious Disease, St. Mary's Campus, Imperial College, London, United Kingdom

Ian M. Jones

Affiliation School of Biological Sciences, University of Reading, Reading, United Kingdom

Ray J. Pickles

Affiliation Cystic Fibrosis/Pulmonary Research and Treatment Center, Department of Microbiology and Immunology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Catherine I. Thompson

Affiliation Virus Reference Department, Health Protection Agency Microbiology Services Colindale, London, United Kingdom

Wendy S. Barclay

w.barclay@imperial.ac.uk

Affiliation Department of Virology, Division of Infectious Disease, St. Mary's Campus, Imperial College, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NvD HS KLR WSB. Performed the experiments: NvD HS KLR. Analyzed the data: NvD HS KLR WSB. Contributed reagents/materials/analysis tools: NvD IMJ RJP CIT. Wrote the paper: NvD HS KLR CIT WSB.