About the Authors

Dean Thumkeo

Affiliation Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto, Japan

Ryota Shinohara

Affiliation Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto, Japan

Keisuke Watanabe

Affiliation Department of Morphological Neural Science, Graduate School of Medical Sciences, Kumamoto University, Kumamoto, Japan

Hirohide Takebayashi

Affiliation Department of Morphological Neural Science, Graduate School of Medical Sciences, Kumamoto University, Kumamoto, Japan

Yosuke Toyoda

Affiliation Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto, Japan

Kiyoshi Tohyama

Affiliation Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto, Japan

Toshimasa Ishizaki

Affiliation Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto, Japan

Tomoyuki Furuyashiki

Affiliation Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto, Japan

Shuh Narumiya

snaru@mfour.med.kyoto-u.ac.jp

Affiliation Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DT SN. Performed the experiments: DT. Analyzed the data: DT SN. Contributed reagents/materials/analysis tools: RS KW HT YT KT TF TI. Wrote the paper: DT TF TI SN. Contributed equally to this work: DT RS.