About the Authors

Akihiro Nasu

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Hiroyuki Marusawa

maru@kuhp.kyoto-u.ac.jp

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yoshihide Ueda

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Norihiro Nishijima

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Ken Takahashi

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yukio Osaki

Affiliation Department of Gastroenterology and Hepatology, Osaka Red Cross Hospital, Osaka, Japan

Yukitaka Yamashita

Affiliation Department of Gastroenterology and Hepatology, Wakayama Red Cross Hospital, Wakayama, Japan

Tetsuro Inokuma

Affiliation Department of Gastroenterology, Kobe City Medical Center General Hospital, Kobe, Japan

Takashi Tamada

Affiliation Department of Gastroenterology and Hepatology Takatsuki Red Cross Hospital, Osaka, Japan

Takeshi Fujiwara

Affiliation Department of Nanobio Drug Discovery, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Fumiaki Sato

Affiliation Department of Nanobio Drug Discovery, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Kazuharu Shimizu

Affiliation Department of Nanobio Drug Discovery, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Tsutomu Chiba

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AN HM. Performed the experiments: AN HM. Analyzed the data: AN HM NN TF FS KS TC YU. Contributed reagents/materials/analysis tools: AN HM YO YY TI TT. Wrote the paper: AN HM KT TC.