About the Authors

Marian C. Aldhous

maldhous@ed.ac.uk

Affiliation Gastrointestinal Unit, Molecular Medicine Centre, Western General Hospital, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Kimberley Soo

Affiliation Gastrointestinal Unit, Molecular Medicine Centre, Western General Hospital, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Lesley A. Stark

Affiliation Edinburgh Cancer Centre and MRC Human Genetics Unit, Institute of Genetics and Molecular Medicine, Western General Hospital, Edinburgh, Scotland, United Kingdom

Agata A. Ulanicka

Affiliation Gastrointestinal Unit, Molecular Medicine Centre, Western General Hospital, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Jennifer E. Easterbrook

Affiliation Gastrointestinal Unit, Molecular Medicine Centre, Western General Hospital, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Malcolm G. Dunlop

Affiliation Edinburgh Cancer Centre and MRC Human Genetics Unit, Institute of Genetics and Molecular Medicine, Western General Hospital, Edinburgh, Scotland, United Kingdom

Jack Satsangi

Affiliation Gastrointestinal Unit, Molecular Medicine Centre, Western General Hospital, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MCA LAS. Performed the experiments: MCA KS AAU JEE. Analyzed the data: MCA KS AAU JEE. Contributed reagents/materials/analysis tools: MCA LAS MGD. Wrote the paper: MCA LAS MGD JS.