About the Authors

Todd W. Arnold

arnol065@umn.edu

Affiliation Department of Fisheries, Wildlife, and Conservation Biology, University of Minnesota, St. Paul, Minnesota, United States of America

Robert M. Zink

Affiliation Bell Museum and Department of Ecology, Evolution and Behavior, University of Minnesota, St. Paul, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TWA RMZ. Analyzed the data: TWA. Wrote the paper: TWA RMZ.