About the Authors

Douglas S. Reed

dsreed@pitt.edu

Affiliation Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Le'Kneitah Smith

Affiliation Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Tammy Dunsmore

Affiliation Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Anita Trichel

Affiliation Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Luis A. Ortiz

Affiliation Department of Environmental & Occupational Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Kelly Stefano Cole

Affiliation Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Eileen Barry

Affiliation Center for Vaccine Development, University of Maryland, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DSR KSC EB. Performed the experiments: DSR LS TD AT. Analyzed the data: DSR LAO AT EB. Wrote the paper: DSR LAO EB.