About the Authors

Taiju Saito

taiju@hokudai-nanae.jp

Affiliations Nanae Fresh Water Laboratory, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan, Laboratory of Aquaculture Genetics & Genomics, Faculty of Fisheries Sciences, Hokkaido University, Hakodate, Japan

Rie Goto-Kazeto

Affiliation Nanae Fresh Water Laboratory, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan

Yutaka Kawakami

Affiliation Nanae Fresh Water Laboratory, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan

Kazuharu Nomura

Affiliation National Research Institute of Aquaculture, 22-1 Nakatsuhamaura, Minami-ise, Mie, Japan

Hideki Tanaka

Affiliation National Research Institute of Aquaculture, 22-1 Nakatsuhamaura, Minami-ise, Mie, Japan

Shinji Adachi

Affiliation Laboratory of Aquaculture Biology, Faculty of Fisheries Sciences, Hokkaido University, Hakodate, Japan

Katsutoshi Arai

Affiliation Laboratory of Aquaculture Genetics & Genomics, Faculty of Fisheries Sciences, Hokkaido University, Hakodate, Japan

Etsuro Yamaha

Affiliation Nanae Fresh Water Laboratory, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TS RG-K KA EY. Performed the experiments: TS RG-K YK. Analyzed the data: TS RG-K. Contributed reagents/materials/analysis tools: KN HT SA. Wrote the paper: TS.