About the Authors

Michael F. Westphal

mwestpha@blm.gov

Affiliation Division of Biology and The Ecological Genomics Institute, Kansas State University, Manhattan, Kansas, United States of America

Jodi L. Massie

Affiliation Department of Biology, Black Hills State University, Spearfish, South Dakota, United States of America

Joanna M. Bronkema

Affiliation Earlham College, Richmond, Indiana, United States of America

Brian E. Smith

Affiliation Department of Biology, Black Hills State University, Spearfish, South Dakota, United States of America

Theodore J. Morgan

Affiliation Division of Biology and The Ecological Genomics Institute, Kansas State University, Manhattan, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MFW JLM BES TJM. Performed the experiments: MFW JLM JMB BES. Analyzed the data: MFW JLM BES TJM. Contributed reagents/materials/analysis tools: MFW JLM BES TJM. Wrote the paper: MFW TJM.