About the Authors

Marguerite R. Irvin

Contributed equally to this work with: Marguerite R. Irvin, Nathan E. Wineinger, Treva K. Rice

irvinr@uab.edu

Affiliation Department of Epidemiology, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Nathan E. Wineinger

Contributed equally to this work with: Marguerite R. Irvin, Nathan E. Wineinger, Treva K. Rice

Affiliation Department of Biostatistics, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Treva K. Rice

Contributed equally to this work with: Marguerite R. Irvin, Nathan E. Wineinger, Treva K. Rice

Affiliation Division of Biostatistics, Washington University, St. Louis, Missouri, United States of America

Nicholas M. Pajewski

Affiliation Department of Biostatistical Sciences, Wake Forest School of Medicine, Winston Salem, North Carolina, United States of America

Edmond K. Kabagambe

Affiliation Department of Epidemiology, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Charles C. Gu

Affiliation Division of Biostatistics, Washington University, St. Louis, Missouri, United States of America

Jim Pankow

Affiliation Department of Epidemiology, University of Minnesota, St. Paul, Minnesota, United States of America

Kari E. North

Affiliations Department of Epidemiology, University of North Carolina, Chapel Hill, North Carolina, United States of America, Carolina Center for Genome Sciences, University of North Carolina, Chapel Hill, North Carolina, United States of America

Jemma B. Wilk

Affiliation Department of Neurology, Boston University, Boston, Massachusetts, United States of America

Barry I. Freedman

Affiliation Department of Internal Medicine/Nephrology, Wake Forest School of Medicine, Winston Salem, North Carolina, United States of America

Nora Franceschini

Affiliation Department of Epidemiology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Uli Broeckel

Affiliation Department of Medicine, Medical College of Wisconsin, Milwaukee, Wisconsin, United States of America

Hemant K. Tiwari

Affiliation Department of Biostatistics, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Donna K. Arnett

Affiliation Department of Epidemiology, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MRI DKA HKT TKR. Performed the experiments: UB JP. Analyzed the data: MRI NEW NMP CCG. Contributed reagents/materials/analysis tools: TKR EKK KEN JBW BIF NF. Wrote the paper: MRI NEW TKR.