About the Authors

Ivan E. Collier

Affiliation Division of Dermatology, Department of Medicine, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Wesley Legant

Affiliation Department of Bioengineering, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Barry Marmer

Affiliation Division of Dermatology, Department of Medicine, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Olga Lubman

Affiliation Department of Pathology, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Saveez Saffarian

Affiliation Department of Physics, University of Utah, Salt Lake City, Utah, United States of America

Tetsuro Wakatsuki

Affiliation Department of Physiology, Biotechnology and Bioengineering Center, Medical College of Wisconsin, Milwaukee, Wisconsin, United States of America

Elliot Elson

Affiliation Department of Biochemistry and Molecular Biophysics, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Gregory I. Goldberg

goldberg@DOM.wustl.edu

Affiliations Division of Dermatology, Department of Medicine, Washington University School of Medicine, Saint Louis, Missouri, United States of America, Department of Biochemistry and Molecular Biophysics, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IEC WL BM OL SS EE GIG. Performed the experiments: IEC WL BM OL TW GIG. Analyzed the data: IEC WL BM OL GIG. Contributed reagents/materials/analysis tools: IEC WL BM OL SS TW EE GIG. Wrote the paper: IEC WL OL EE GIG.