About the Authors

Herman Tse

Affiliation State Key Laboratory for Emerging Infectious Diseases, Carol Yu Centre for Infection, The University of Hong Kong, Hong Kong, China

Kelvin K. W. To

Affiliation State Key Laboratory for Emerging Infectious Diseases, Carol Yu Centre for Infection, The University of Hong Kong, Hong Kong, China

Xi Wen

Affiliation Department of Microbiology, The University of Hong Kong, Hong Kong, China

Honglin Chen

Affiliation State Key Laboratory for Emerging Infectious Diseases, Carol Yu Centre for Infection, The University of Hong Kong, Hong Kong, China

Kwok-Hung Chan

Affiliation Department of Microbiology, Queen Mary Hospital, Hong Kong, China

Hoi-Wah Tsoi

Affiliation Department of Microbiology, The University of Hong Kong, Hong Kong, China

Iris W. S. Li

Affiliation Department of Microbiology, Queen Mary Hospital, Hong Kong, China

Kwok-Yung Yuen

kyyuen@hkucc.hku.hk

Affiliation State Key Laboratory for Emerging Infectious Diseases, Carol Yu Centre for Infection, The University of Hong Kong, Hong Kong, China

Competing Interests

We gratefully acknowledge funding support from the Providence Foundation Limited in memory of the late Dr Lui Hac Minh, the University Grants Council. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HT KT XW HC K-HC H-WT IL K-YY. Performed the experiments: HT XW HC KHC H-WT. Analyzed the data: HT KT IL K-YY. Contributed reagents/materials/analysis tools: HT KT XW HC K-HC H-WT IL K-YY. Wrote the paper: HT KT IL K-YY.