About the Authors

Qiuling He

Affiliations College of Forest Resources and Environment, Nanjing Forestry University, Nanjing, People's Republic of China, Department of Agronomy, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou, People's Republic of China

Ye Shen

Affiliation College of Forest Resources and Environment, Nanjing Forestry University, Nanjing, People's Republic of China

Mingxiu Wang

Affiliation College of Forest Resources and Environment, Nanjing Forestry University, Nanjing, People's Republic of China

Minren Huang

mrhuang@njfu.edu.cn (MH); wanglsh@ibcas.ac.cn (LW); rwu@hes.hmc.psu.edu (RW)

Affiliation College of Forest Resources and Environment, Nanjing Forestry University, Nanjing, People's Republic of China

Ruizhen Yang

Affiliation Beijing Botanical Garden, Institute of Botany, Chinese Academy of Sciences, Beijing, People's Republic of China

Shuijin Zhu

Affiliation Department of Agronomy, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou, People's Republic of China

Liangsheng Wang

mrhuang@njfu.edu.cn (MH); wanglsh@ibcas.ac.cn (LW); rwu@hes.hmc.psu.edu (RW)

Affiliation Beijing Botanical Garden, Institute of Botany, Chinese Academy of Sciences, Beijing, People's Republic of China

Yanjun Xu

Affiliation Department of Applied Chemistry, China Agricultural University, Beijing, People's Republic of China

Rongling Wu

mrhuang@njfu.edu.cn (MH); wanglsh@ibcas.ac.cn (LW); rwu@hes.hmc.psu.edu (RW)

Affiliations Center for Computational Biology, Beijing Forestry University, Beijing, People's Republic of China, National Engineering Laboratory for Tree Breeding, Beijing Forestry University, Beijing, People's Republic of China, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Beijing Forestry University, Beijing, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MH LW MW. Performed the experiments: QH YS RY SZ YX. Analyzed the data: QH RW. Contributed reagents/materials/analysis tools: QH YX. Wrote the paper: QH RW.