About the Authors

Antony B. Holmes

Affiliations Department of Biomedical Informatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America, Center for Computational Biology and Bioinformatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Alexander Hawson

Affiliations Center for Computational Biology and Bioinformatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America, Department of Medicine, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Feng Liu

Affiliation Department of Biomedical Informatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Carol Friedman

Affiliation Department of Biomedical Informatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Hossein Khiabanian

hossein@c2b2.columbia.edu

Affiliations Department of Biomedical Informatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America, Center for Computational Biology and Bioinformatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Raul Rabadan

Affiliations Department of Biomedical Informatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America, Center for Computational Biology and Bioinformatics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ABH AH CF HK RR. Performed the experiments: ABH AH FL HK. Analyzed the data: ABH AH HK RR. Contributed reagents/materials/analysis tools: ABH AH CF HK RR. Wrote the paper: ABH AH CF HK RR.