About the Authors

Shaukat Ali

Affiliation Institute of Biology, Sylvius Laboratory, Leiden University, Leiden, The Netherlands

Harald G. J. van Mil

Affiliations Institute of Biology, Sylvius Laboratory, Leiden University, Leiden, The Netherlands, Mathematical Institute, Leiden University, Leiden, The Netherlands

Michael K. Richardson

m.k.richardson@biology.leidenuniv.nl

Affiliation Institute of Biology, Sylvius Laboratory, Leiden University, Leiden, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SA MKR HGJvM. Performed the experiments: SA. Analyzed the data: SA MKR HGJvM. Contributed reagents/materials/analysis tools: MKR. Wrote the paper: SA MKR.