About the Authors

Nao Horio

Contributed equally to this work with: Nao Horio, Ryusuke Yoshida

Affiliation Section of Oral Neuroscience, Graduate School of Dental Science, Kyushu University, Fukuoka, Japan

Ryusuke Yoshida

Contributed equally to this work with: Nao Horio, Ryusuke Yoshida

Affiliation Section of Oral Neuroscience, Graduate School of Dental Science, Kyushu University, Fukuoka, Japan

Keiko Yasumatsu

Affiliations Section of Oral Neuroscience, Graduate School of Dental Science, Kyushu University, Fukuoka, Japan, Department of Oral Physiology, Asahi University School of Dentistry, Gifu, Japan

Yuchio Yanagawa

Affiliations Department of Genetic and Behavioral Neuroscience, Gunma University Graduate School of Medicine, Maebashi, Japan, JST, CREST, Tokyo, Japan

Yoshiro Ishimaru

Affiliations Department of Applied Biological Chemistry, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan, Department of Molecular Genetics and Microbiology, Duke University Medical Center, Durham, North Carolina, United States of America

Hiroaki Matsunami

Affiliation Department of Molecular Genetics and Microbiology, Duke University Medical Center, Durham, North Carolina, United States of America

Yuzo Ninomiya

yuninom@dent.kyushu-u.ac.jp

Affiliation Section of Oral Neuroscience, Graduate School of Dental Science, Kyushu University, Fukuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN. Performed the experiments: NH RY KY YI. Analyzed the data: NH RY HM YN. Contributed reagents/materials/analysis tools: NH RY YI YY. Wrote the paper: NH RY HM YN.