About the Authors

Lydia Beaudrot

Affiliations Graduate Group in Ecology, University of California Davis, Davis, California, United States of America, International Field Biology Course 2008, Arnold Arboretum, Center for Tropical Forest Science, Pasoh Forest Reserve, Malaysia

Yanjun Du

Affiliations International Field Biology Course 2008, Arnold Arboretum, Center for Tropical Forest Science, Pasoh Forest Reserve, Malaysia, Institute of Botany, Chinese Academy of Sciences, Beijing, China

Abdul Rahman Kassim

Affiliations International Field Biology Course 2008, Arnold Arboretum, Center for Tropical Forest Science, Pasoh Forest Reserve, Malaysia, Forest Research Institute Malaysia, Selangor, Malaysia

Marcel Rejmánek

Affiliations Graduate Group in Ecology, University of California Davis, Davis, California, United States of America, Department of Evolution and Ecology, University of California Davis, Davis, California, United States of America

Rhett D. Harrison

rharrison@xtbg.org.cn

Affiliations International Field Biology Course 2008, Arnold Arboretum, Center for Tropical Forest Science, Pasoh Forest Reserve, Malaysia, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan Province, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LB YD RDH. Performed the experiments: LB YD. Analyzed the data: LB YD MR. Contributed reagents/materials/analysis tools: ARK. Wrote the paper: LB YD RDH. Commented on a draft of the manuscript: LB YD ARK MR RDH.