About the Authors

Yoshihiro Omori

Contributed equally to this work with: Yoshihiro Omori, Kimiko Katoh, Shigeru Sato

Affiliations Department of Developmental Biology, Osaka Bioscience Institute, Osaka, Japan, JST, CREST, Osaka, Japan, PRESTO, Suita, Osaka, Japan

Kimiko Katoh

Contributed equally to this work with: Yoshihiro Omori, Kimiko Katoh, Shigeru Sato

Affiliation Department of Developmental Biology, Osaka Bioscience Institute, Osaka, Japan

Shigeru Sato

Contributed equally to this work with: Yoshihiro Omori, Kimiko Katoh, Shigeru Sato

Affiliations Department of Developmental Biology, Osaka Bioscience Institute, Osaka, Japan, Department of Ophthalmology, Osaka University Graduate School of Medicine, Suita, Osaka, Japan, Department of Visual Science, Osaka University Graduate School of Medicine, Suita, Osaka, Japan

Yuki Muranishi

Affiliation Department of Developmental Biology, Osaka Bioscience Institute, Osaka, Japan

Taro Chaya

Affiliations Department of Developmental Biology, Osaka Bioscience Institute, Osaka, Japan, JST, CREST, Osaka, Japan

Akishi Onishi

Affiliations Department of Developmental Biology, Osaka Bioscience Institute, Osaka, Japan, JST, CREST, Osaka, Japan

Takashi Minami

Affiliation The Research Center for Advanced Science and Technology, The University of Tokyo, Tokyo, Japan

Takashi Fujikado

Affiliations Department of Ophthalmology, Osaka University Graduate School of Medicine, Suita, Osaka, Japan, Department of Visual Science, Osaka University Graduate School of Medicine, Suita, Osaka, Japan

Takahisa Furukawa

furukawa@obi.or.jp

Affiliations Department of Developmental Biology, Osaka Bioscience Institute, Osaka, Japan, JST, CREST, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: T. Furukawa. Performed the experiments: YO KK SS YM TC AO TM. Analyzed the data: YO KK SS T. Furukawa. Wrote the paper: YO T. Furukawa. Supervised the experiments: T. Fujikado T. Furukawa.