About the Authors

Shiva Kumar

Affiliations Laboratory of Structural Sciences, Van Andel Research Institute, Grand Rapids, Michigan, United States of America, Department of Biological Sciences, National University of Singapore, Singapore

Augen Pioszak

Current address: Department of Biochemistry and Molecular Biology, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America

Affiliation Laboratory of Structural Sciences, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Chenghai Zhang

Affiliation Laboratory of Structural Sciences, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Kunchithapadam Swaminathan

eric.xu@vai.org (HEX); dbsks@nus.edu.sg (KS)

Affiliation Department of Biological Sciences, National University of Singapore, Singapore

H. Eric Xu

eric.xu@vai.org (HEX); dbsks@nus.edu.sg (KS)

Affiliation Laboratory of Structural Sciences, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Performed the experiments: SK. Conceived the project: HEX AP. Supervised the project: HEX AP KS. Helped set up CRE-luciferase reporter experiments: CZ. Helped in structure determination: AP KS.