About the Authors

Yuji Hasegawa


Affiliation Honda Research Institute Japan Co. Ltd., Wako-shi, Saitama, Japan

Hidetoshi Ikeno

Affiliation School of Human Science and Environment, University of Hyogo, Honmachi, Shinzaike, Himeji-shi, Japan

Competing Interests

YH is employed by a commercial company (Honda Research Institute Japan Co. Ltd). YH confirmed that this does not alter his adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YH. Performed the experiments: YH HI. Analyzed the data: YH. Contributed reagents/materials/analysis tools: YH HI. Wrote the manuscript: YH HI.