About the Authors

Dongxing Zhu

Affiliation The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, Scotland, United Kingdom

Neil Charles Wallace Mackenzie

Affiliation The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, Scotland, United Kingdom

José Luis Millán

Affiliation Sanford Children's Health Research Center, Sanford-Burnham Medical Research Institute, La Jolla, California, United States of America

Colin Farquharson

Affiliation The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, Scotland, United Kingdom

Vicky Elizabeth MacRae

vicky.macrae@roslin.ed.ac.uk

Affiliation The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JLM CF VEM. Performed the experiments: DZ NCWM VEM. Analyzed the data: DZ VEM. Contributed reagents/materials/analysis tools: JLM VEM. Wrote the paper: DZ NCWM JLM CF VEM.