About the Authors

Kodzue Kinoshita

Affiliation Faunal Diversity Sciences, Graduate School of Agriculture, Kobe University, Nada-ku, Kobe, Japan

Sayaka Inada

Affiliation Faunal Diversity Sciences, Graduate School of Agriculture, Kobe University, Nada-ku, Kobe, Japan

Kazuya Seki

Affiliation Kobe Municipal Oji Zoo, Nada-ku, Kobe, Japan

Aiko Sasaki

Affiliation Tama Zoological Park, Hino-shi, Tokyo, Japan

Natsuki Hama

Current address: Kobe Institute of Health, Chuo-ku, Kobe, Japan

Affiliation Kobe Municipal Oji Zoo, Nada-ku, Kobe, Japan

Hiroshi Kusunoki

kusunoki@kobe-u.ac.jp

Affiliation Faunal Diversity Sciences, Graduate School of Agriculture, Kobe University, Nada-ku, Kobe, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HK. Performed the experiments: KK SI. Analyzed the data: KK. Contributed reagents/materials/analysis tools: NH KS AS. Wrote the paper: KK.