About the Authors

Shinjae Chung

Contributed equally to this work with: Shinjae Chung, Michel M. M. Verheij

Affiliation Department of Pharmacology, University of California Irvine, Irvine, California, United States of America

Michel M. M. Verheij

Contributed equally to this work with: Shinjae Chung, Michel M. M. Verheij

Affiliation Department of Molecular Animal Physiology, Donders Institute for Brain, Cognition and Behaviour, and Nijmegen Center for Molecular Life Sciences, Radboud University, Nijmegen, The Netherlands

Peter Hesseling

Affiliation Department of Molecular Animal Physiology, Donders Institute for Brain, Cognition and Behaviour, and Nijmegen Center for Molecular Life Sciences, Radboud University, Nijmegen, The Netherlands

Ruben W. M. van Vugt

Affiliation Department of Molecular Animal Physiology, Donders Institute for Brain, Cognition and Behaviour, and Nijmegen Center for Molecular Life Sciences, Radboud University, Nijmegen, The Netherlands

Mahalah Buell

Affiliation Department of Psychiatry, University of California San Diego, La Jolla, California, United States of America

James D. Belluzzi

Affiliation Department of Pharmacology, University of California Irvine, Irvine, California, United States of America

Mark A. Geyer

Affiliation Department of Psychiatry, University of California San Diego, La Jolla, California, United States of America

Gerard J. M. Martens

Affiliation Department of Molecular Animal Physiology, Donders Institute for Brain, Cognition and Behaviour, and Nijmegen Center for Molecular Life Sciences, Radboud University, Nijmegen, The Netherlands

Olivier Civelli

ocivelli@uci.edu

Affiliations Department of Pharmacology, University of California Irvine, Irvine, California, United States of America, Department of Developmental and Cell Biology, University of California Irvine, Irvine, California, United States of America, Department of Pharmaceutical Sciences, University of California Irvine, Irvine, California, United States of America

Competing Interests

No authors have any competing financial interests in relation to the work described. MG holds an equity interest in San Diego Instruments, Inc. This does not alter the authors adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SC OC. Performed the experiments: SC MV PH RvV. Analyzed the data: SC MV MB MG GM OC JB. Contributed reagents/materials/analysis tools: MB MG. Wrote the paper: SC MV OC.