About the Authors

René A. W. Frank

Contributed equally to this work with: René A. W. Frank, Allan F. McRae

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Allan F. McRae

Contributed equally to this work with: René A. W. Frank, Allan F. McRae

Affiliation Queensland Institute of Medical Research, Royal Brisbane Hospital, Brisbane, Australia

Andrew J. Pocklington

Affiliation School of Informatics, Edinburgh University, Edinburgh, United Kingdom

Louie N. van de Lagemaat

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Pau Navarro

Affiliation MRC Human Genetics, Institute of Genetics and Molecular Medicine, Western General Hospital, Edinburgh, United Kingdom

Mike D. R. Croning

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Noboru H. Komiyama

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Sophie J. Bradley

Affiliation Department of Cell Physiology and Pharmacology, University of Leicester, Leicester, United Kingdom

R. A. John Challiss

Affiliation Department of Cell Physiology and Pharmacology, University of Leicester, Leicester, United Kingdom

J. Douglas Armstrong

Affiliation School of Informatics, Edinburgh University, Edinburgh, United Kingdom

Robert D. Finn

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Mary P. Malloy

Affiliation Division of Psychiatry, University of Edinburgh, Royal Edinburgh Hospital, Edinburgh, United Kingdom

Alan W. MacLean

Affiliation Division of Psychiatry, University of Edinburgh, Royal Edinburgh Hospital, Edinburgh, United Kingdom

Sarah E. Harris

Affiliation Department of Psychology, Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom

John M. Starr

Affiliation Department of Psychology, Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom

Sanjeev S. Bhaskar

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Eleanor K. Howard

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Sarah E. Hunt

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Alison J. Coffey

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Venkatesh Ranganath

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Panos Deloukas

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Jane Rogers

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Walter J. Muir

Affiliation Division of Psychiatry, University of Edinburgh, Royal Edinburgh Hospital, Edinburgh, United Kingdom

Ian J. Deary

Affiliation Department of Psychology, Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom

Douglas H. Blackwood

Affiliation Division of Psychiatry, University of Edinburgh, Royal Edinburgh Hospital, Edinburgh, United Kingdom

Peter M. Visscher

Affiliation Queensland Institute of Medical Research, Royal Brisbane Hospital, Brisbane, Australia

Seth G. N. Grant

sg3@sanger.ac.uk

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SGNG PMV DHB IJD WJM JR NHK RAWF RAJC. Performed the experiments: RAWF AFM SJB SEH JMS SSB EKH VR PD MPM. Analyzed the data: RAWF AFM AJP LNdvL PN MDRC SJB RAJC JDA RDF SSB SEH AJC VR PD IJD PMV SGNG. Contributed reagents/materials/analysis tools: SGNG PMV DHB IJD WJM JR PD RAJC. Wrote the paper: RAWF AFM LNdvL AJP IJD DHB PMV SGNG.