About the Authors

M. Cecilia Cirio

Affiliation Department of Developmental Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Zhao Hui

Affiliation School of Biomedical Sciences, The Chinese University of Hong Kong, Ma Liu Shui, Hong Kong

Caroline E. Haldin

Affiliation Department of Developmental Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Chiara Cianciolo Cosentino

Affiliation Department of Developmental Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Carsten Stuckenholz

Affiliation Department of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Xiongfong Chen

Affiliation Unit on Biologic Computation, Eunice Kennedy Shriver National Institute of Child Health and Human Development, National Institutes of Health (NIH), Bethesda, Maryland, United States of America

Sung-Kook Hong

Affiliation Laboratory of Molecular Genetics, Eunice Kennedy Shriver National Institute of Child Health and Human Development, National Institutes of Health (NIH), Bethesda, Maryland, United States of America

Igor B. Dawid

Affiliation Laboratory of Molecular Genetics, Eunice Kennedy Shriver National Institute of Child Health and Human Development, National Institutes of Health (NIH), Bethesda, Maryland, United States of America

Neil A. Hukriede

hukriede@pitt.edu

Affiliations Department of Developmental Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Drug Discovery Institute, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MCC IBD NAH. Performed the experiments: MCC ZH CH CCC XC SKH NAH. Analyzed the data: MCC ZH CH CCC CS XC SKH IBD NAH. Contributed reagents/materials/analysis tools: CS XC IBD NAH. Wrote the paper: MCC NAH.