About the Authors

Yuan Yuan Shi

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Zachary Y. Huang

bees1965@sina.com (ZJZ); bees@msu.edu (ZYH)

Affiliations Department of Entomology, Michigan State University, East Lansing, Michigan, United States of America, Ecology, Evolutionary Biology and Behavior Program, Michigan State University, East Lansing, Michigan, United States of America

Zhi Jiang Zeng

bees1965@sina.com (ZJZ); bees@msu.edu (ZYH)

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Zi Long Wang

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Xiao Bo Wu

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Wei Yu Yan

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YYS ZJZ ZLW ZYH. Performed the experiments: YYS XBW WYY. Analyzed the data: ZYH YYS. Contributed reagents/materials/analysis tools: ZJZ. Wrote the paper: ZYH YYS ZJZ. All authors approved the manuscript.