About the Authors

Thomas Owen Richardson

tom.richardson@bristol.ac.uk

Affiliations Department of Engineering, Design and Mathematics, University of the West of England, Bristol, United Kingdom, School of Biological Sciences, University of Bristol, Bristol, United Kingdom

Kim Christensen

Affiliations Institute for Mathematical Sciences, Imperial College London, London, United Kingdom, Department of Physics, Imperial College London, London, United Kingdom

Nigel Rigby Franks

Affiliation School of Biological Sciences, University of Bristol, Bristol, United Kingdom

Henrik Jeldtoft Jensen

Affiliations Institute for Mathematical Sciences, Imperial College London, London, United Kingdom, Department of Mathematics, Imperial College London, London, United Kingdom

Ana Blagovestova Sendova-Franks

Affiliation Department of Engineering, Design and Mathematics, University of the West of England, Bristol, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TOR ABSF NRF KC HJJ. Performed the experiments: TOR. Analyzed the data: TOR. Contributed reagents/materials/analysis tools: TOR ABSF HJJ KC. Wrote the paper: TOR. Provided feedback on the modelling: ABSF, KC, HJJ, NRF. Provided feedback on the manuscript: ABSF, KC, HJJ, NRF.