About the Authors

Harri Lempiäinen

Contributed equally to this work with: Harri Lempiäinen, Arne Müller

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Arne Müller

Contributed equally to this work with: Harri Lempiäinen, Arne Müller

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Sarah Brasa

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Soon-Siong Teo

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Tim-Christoph Roloff

Affiliation Friedrich Miescher Institute for Biomedical Research, Basel, Switzerland

Laurent Morawiec

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Natasa Zamurovic

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Axel Vicart

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Enrico Funhoff

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Philippe Couttet

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Dirk Schübeler

Affiliation Friedrich Miescher Institute for Biomedical Research, Basel, Switzerland

Olivier Grenet

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Jennifer Marlowe

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Jonathan Moggs

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Rémi Terranova

remi.terranova@novartis.com

Affiliation Investigative Toxicology, Preclinical Safety, Translational Sciences, Novartis Institutes for Biomedical Research, Basel, Switzerland

Competing Interests

The authors have declared that no competing interests exist. All authors work for Novartis Institutes for Biomedical Research and confirm that this does not alter their adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: J. Marlowe J. Moggs OG RT HL EF NZ. Performed the experiments: HL SB S-ST LM AV. Analyzed the data: HL AM SB S-ST T-CR AV EF PC DS OG J. Marlowe J. Moggs RT. Contributed reagents/materials/analysis tools: T-CR DS. Wrote the paper: RT AM EF HL J. Marlowe J. Moggs.