About the Authors

Naomi J. Fox

Affiliations Animal Health, Scottish Agricultural College, Edinburgh, United Kingdom, Environment Department, University of York, York, United Kingdom, Biomathematics and Statistics Scotland, Edinburgh, United Kingdom

Piran C. L. White

Affiliation Environment Department, University of York, York, United Kingdom

Colin J. McClean

Affiliation Environment Department, University of York, York, United Kingdom

Glenn Marion

Affiliation Biomathematics and Statistics Scotland, Edinburgh, United Kingdom

Andy Evans

Affiliation Animal Health, Scottish Agricultural College, Edinburgh, United Kingdom

Michael R. Hutchings

mike.hutchings@sac.ac.uk

Affiliation Animal Health, Scottish Agricultural College, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NJF MRH PCLW CJM AE GM. Performed the experiments: NJF CJM. Analyzed the data: NJF. Wrote the paper: NJF MRH PCLW GM.