About the Authors

Hiroki Obata

totori@kumamoto-u.ac.jp

Affiliation Faculty of Letters, Kumamoto University, Kumamoto City, Kumamoto Prefecture, Japan

Aya Manabe

Affiliation Faculty of Letters, Kumamoto University, Kumamoto City, Kumamoto Prefecture, Japan

Naoko Nakamura

Affiliation Faculty of Law and Letters, Kagoshima University, Kagoshima City, Kagoshima Prefecture, Japan

Tomokazu Onishi

Affiliation Faculty of Intercultural Studies, The International University of Kagoshima, Kagoshima City, Kagoshima Prefecture, Japan

Yasuko Senba

Affiliation Project for Seed Impression Studies, Kumamoto University, Kumamoto City, Kumamoto Prefecture, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: HO. Performed lab work: AM. Took SEM Photos: AM. Co-Searcher of the findings: AM NN TO YS.