About the Authors

Shannon Balfry

balfry@mail.ubc.ca

Current address: DFO/UBC Centre for Aquaculture and Environmental Research, West Vancouver, British Columbia, Canada

Affiliation Faculty of Land and Food Systems, The University of British Columbia, Vancouver, British Columbia, Canada

David W. Welch

Affiliation Kintama Research Services Ltd., Nanaimo, British Columbia, Canada

Jody Atkinson

Affiliation Department of Zoology, The University of British Columbia, Vancouver, British Columbia, Canada

Al Lill

Affiliation Living Rivers–Georgia Basin/Vancouver Island, North Vancouver, British Columbia, Canada

Stephen Vincent

Affiliation Seymour Salmonid Society, North Vancouver, British Columbia, Canada

Competing Interests

Two of the co-authors (Welch and Lill) were employed by commercial companies (Kintama Research Corporation and Living Rivers) that provided their time/salary (in-kind funding), which enabled their participation in this research project. The involvement of these co-authors and their respective association with commercial companies does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SB DWW AL SV. Performed the experiments: SB DWW JA SV. Analyzed the data: SB DWW JA AL SV. Contributed reagents/materials/analysis tools: SB DWW JA SV. Wrote the paper: SB DWW JA AL SV.