About the Authors

Alan Yung-Chih Hu

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Yu-Fen Tseng

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Tsai-Chuan Weng

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Chien-Chun Liao

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Johnson Wu

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Ai-Hsiang Chou

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Hsin-Ju Chao

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Anna Gu

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Janice Chen

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Su-Chen Lin

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Chia-Hsin Hsiao

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority

Suh-Chin Wu

Affiliations Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority, Institute of Biotechnology, National Tsing Hua University, Hsinchu, Taiwan Authority

Pele Chong

pelechong@nhri.org.tw

Affiliations Vaccine Research and Development Center, National Health Research Institutes, Zhunan, Taiwan Authority, Graduate Institute of Immunology, China Medical University, Taichung, Taiwan Authority

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AYCH PC. Performed the experiments: AYCH YFT TCW CCL JW HJC AG JC SCL. Analyzed the data: AYCH AHC CHH SCW PC. Contributed reagents/materials/analysis tools: AHC CHH. Wrote the paper: AYCH PC. Secured funding from Taiwanese Government: PC.