About the Authors

Yuanquan Song

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jason R. Willer

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Paul C. Scherer

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jessica A. Panzer

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Amy Kugath

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Emmanuel Skordalakes

Affiliation Wistar Institute, Philadelphia, Pennsylvania, United States of America

Ronald G. Gregg

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Gregory B. Willer

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Rita J. Balice-Gordon

rbaliceg@mail.med.upenn.edu

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YS RBG. Performed the experiments: YS JRW PCS JAP AK RGG GBW. Analyzed the data: YS ES RBG. Wrote the paper: YS RBG.