About the Authors
- Hirokata Fukushima
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* E-mail: hirof@ardbeg.c.u-tokyo.ac.jp (HF); khiraki@idea.c.u-tokyo.ac.jp (KH)
Affiliations Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan, Japan Society for Promotion of Sciences, Tokyo, Japan, Department of Psychology, Keio University, Tokyo, Japan
- Satoshi Hirata
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Affiliation Great Ape Research Institute of Hayashibara Biochemical Laboratories, Inc., Tamano, Japan
- Ari Ueno
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Affiliations Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan, Department of Human Relations Studies, School of Human Cultures, The University of Shiga Prefecture, Hikone, Japan
- Goh Matsuda
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Affiliation Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan
- Kohki Fuwa
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Affiliation Great Ape Research Institute of Hayashibara Biochemical Laboratories, Inc., Tamano, Japan
- Keiko Sugama
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Affiliation Great Ape Research Institute of Hayashibara Biochemical Laboratories, Inc., Tamano, Japan
- Kiyo Kusunoki
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Affiliation Great Ape Research Institute of Hayashibara Biochemical Laboratories, Inc., Tamano, Japan
- Masahiro Hirai
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Affiliations Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan, Japan Society for Promotion of Sciences, Tokyo, Japan, Department of Psychology, Queen's University, Kingston, Ontario, Canada
- Kazuo Hiraki
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* E-mail: hirof@ardbeg.c.u-tokyo.ac.jp (HF); khiraki@idea.c.u-tokyo.ac.jp (KH)
Affiliation Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan
- Masaki Tomonaga
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Affiliation Section of Language and Intelligence, Primate Research Institute, Kyoto University, Inuyama, Japan
- Toshikazu Hasegawa
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Affiliation Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan
Competing Interests
Four authors are employed by Hayashibara Biochemical Laboratories, Inc. However, the company had no role in study design, consultancy, data collection and analysis, decision to publish, or preparation of the manuscript. The authors declare their full adherence to all PLoS ONE policies on data sharing and materials.
Author Contributions
Conceived and designed the experiments: HF SH AU GM MH KH MT TH. Performed the experiments: SH KF KS KK. Analyzed the data: HF. Contributed reagents/materials/analysis tools: HF GM. Wrote the paper: HF SH AU GM MH KH.