About the Authors

Céline H. Frère

c.frere@uq.edu.au

Affiliation School of Veterinary Sciences, University of Queensland, Brisbane, Australia

Ewa Krzyszczyk

Affiliation Departments of Biology and Psychology, Georgetown University, Washington, District of Columbia, United States of America

Eric M. Patterson

Affiliation Departments of Biology and Psychology, Georgetown University, Washington, District of Columbia, United States of America

Sue Hunter

Affiliation National Aquarium, Baltimore, Maryland, United States of America

Alison Ginsburg

Affiliation National Aquarium, Baltimore, Maryland, United States of America

Janet Mann

Affiliation Departments of Biology and Psychology, Georgetown University, Washington, District of Columbia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CHF EK EMP JM. Performed the experiments: CHF EK EMP SH AG JM. Analyzed the data: CHF. Contributed reagents/materials/analysis tools: CHF SH AG JM. Wrote the paper: CHF EK EMP JM.