About the Authors

Lisette A. H. M. Cornelissen

Affiliation Central Veterinary Institute of Wageningen University, Lelystad, The Netherlands

Robert P. de Vries

Affiliation Virology Division, Department of Infectious Diseases & Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Els A. de Boer-Luijtze

Affiliation Central Veterinary Institute of Wageningen University, Lelystad, The Netherlands

Alan Rigter

Affiliation Virology Division, Department of Infectious Diseases & Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Peter J. M. Rottier

Affiliation Virology Division, Department of Infectious Diseases & Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Cornelis A. M. de Haan

c.a.m.dehaan@uu.nl

Affiliation Virology Division, Department of Infectious Diseases & Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Competing Interests

The authors P.J.M.R. and C.A.M.d.H. have filed a patent application based on findings reported in this study. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LC RdV PJMR CAMdH. Performed the experiments: LC RdV EAdBL AR. Analyzed the data: LC RdV CAMdH. Wrote the paper: LC PJMR CAMdH.