About the Authors

Susan M. Twine

susan.twine@nrc-cnrc.gc.ca

Affiliation National Research Council Institute for Biological Sciences, Ottawa, Ontario, Canada

Mireille D. Petit

Affiliation National Research Council Institute for Biological Sciences, Ottawa, Ontario, Canada

Kelly M. Fulton

Affiliation National Research Council Institute for Biological Sciences, Ottawa, Ontario, Canada

Robert V. House

Affiliation DynPort Vaccine Company, LLC, Frederick, Maryland, United States of America

J. Wayne Conlan

Affiliation National Research Council Institute for Biological Sciences, Ottawa, Ontario, Canada

Competing Interests

One listed author is an employee of the Dynport Vaccine Company, a commerical entity. This company administered a contract from the National Institutes of Health, as detailed in the financial disclosure. The company was not a sponsor of this study and the author had no financial contribution outside the federally awarded funds. The authors, therefore, do not believe there to be any competing interests. This does not alter their adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SMT RVH JWC. Performed the experiments: MDP KMF. Analyzed the data: SMT KMF. Contributed reagents/materials/analysis tools: SMT MDP. Wrote the paper: SMT JWC.