About the Authors

Alan D. Pemberton

alan.pemberton@ed.ac.uk

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, Midlothian, United Kingdom

Jeremy K. Brown

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, Midlothian, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AP. Performed the experiments: AP JKB. Analyzed the data: AP JKB. Wrote the paper: AP.