About the Authors

Keizo Takao

Affiliations Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan, Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan, Japan Science and Technology Agency (JST), Core Research for Evolutional Science and Technology (CREST), Kawaguchi, Japan

Koichi Tanda

Affiliation Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan

Kenji Nakamura

Affiliation Mouse Genome Technology Laboratory, Mitsubishi Kagaku Institute of Life Sciences, Machida, Tokyo, Japan

Jiro Kasahara

Affiliations Department of Neurobiology and Therapeutics, Graduate School of Health Bioscience, The University of Tokushima, Tokushima, Japan, Department of Pharmacology, Graduate School of Pharmaceutical Sciences, Tohoku University, Sendai, Japan

Kazuki Nakao

Affiliation Laboratory for Animal Resources and Genetic Engineering, Center for Developmental Biology, RIKEN, Chuo-ku, Kobe, Japan

Motoya Katsuki

Affiliation National Institutes of Natural Sciences, Toranomon, Minato-ku, Tokyo, Japan

Kazuo Nakanishi

Affiliation Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan

Nobuyuki Yamasaki

Affiliation Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan

Keiko Toyama

Affiliations Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan, Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan, Japan Science and Technology Agency (JST), Core Research for Evolutional Science and Technology (CREST), Kawaguchi, Japan

Minami Adachi

Affiliation Department of Neurobiology and Therapeutics, Graduate School of Health Bioscience, The University of Tokushima, Tokushima, Japan

Masahiro Umeda

Affiliation Department of Neurobiology and Therapeutics, Graduate School of Health Bioscience, The University of Tokushima, Tokushima, Japan

Tsutomu Araki

Affiliation Department of Neurobiology and Therapeutics, Graduate School of Health Bioscience, The University of Tokushima, Tokushima, Japan

Kohji Fukunaga

Affiliation Department of Pharmacology, Graduate School of Pharmaceutical Sciences, Tohoku University, Sendai, Japan

Hisatake Kondo

Affiliation Department of Rehabilitation, Faculty of Medical Science & Welfare, Tohoku Bunka Gakuen University, Sendai, Japan

Hiroyuki Sakagami

Affiliation Department of Anatomy, Kitasato University School of Medicine, Sagamihara, Japan

Tsuyoshi Miyakawa

miyakawa@fujita-hu.ac.jp

Affiliations Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan, Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan, Japan Science and Technology Agency (JST), Core Research for Evolutional Science and Technology (CREST), Kawaguchi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TM. Performed the experiments: KT KT JK KN NY KT MA MU TA KF. Analyzed the data: KT KT KN. Contributed reagents/materials/analysis tools: KN KN MK HK HS TM. Wrote the paper: KT KT HS TM.